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- Wisconsin Alumni Research Foundation (WARF) Named Professorship – Vannevar Bush Professor of Biochemistry – 2019
- Member, National Research Committee on Next Generation Researchers – 2016-2018
- Chair, President’s Committee on National Medal of Science committee – 2015-2017
- Vilas Board of Trustees – 2014-present
- Member, President’s Committee on National Medal of Science – 2014-2015
- President’s Committee on National Medal of Science – 2012-2014
- Chair, ASCB International Affairs Committee – 2013-2014
- Adjunct Professor, National Institute of Genetics, Mishima, Japan – 2009-2011
- Council, National Academy of Sciences – 2008-2010
- Fellow, American Association for the Advancement of Science – 2006
- President, Society for Developmental Biology – 2004-2005
- Hilldale Award, Biological Sciences Division, UW-Madison – 2002-2003
- Elected, American Philosophical Society – 2002
- Vilas Professor, UW-Madison – 2001-Present
- President, Genetics Society of America – 2000
- Elected, National Academy of Sciences – 1995
- Elected, American Academy of Arts and Sciences – 1995
- WARF, Mid-Career Award, UW-Madison – 1995-2005
- Investigator, Howard Hughes Medical Institute – 1994
- H.I. Romnes Faculty Fellowship, UW-Madison – 1990-1995
- Pound Award for Excellence in Research, UW-Madison – 1988
- NIH Research Career Development Award – 1984-1989
- NIH Postdoctoral Fellow – 1980-1982
- Jane Coffin Childs Postdoctoral Fellow – 1978-1980
- NSF Predoctoral Fellow, U. Colorado, Boulder – 1974-1977
- Anna Fullam Award, U. Colorado, Boulder – 1973